2. (stative) be hard, not friable (of soil).
He ioio te oneone o te whenua nei (W 1971:78). / The soil of this land is not friable.
3. (stative) be stiff (from being tired).
Ka ioio ōku waewae i te roa o te haerenga (W 1971:78). / My legs were stiff from the long trip.
4. (noun) sinew, muscle, nerve, nervous system.
Ko te roro te pū o te pūnaha ioio e whiwhi ana, e tuku ana i ngā karere, e whakahaere ana i ngā wāhanga katoa o te tinana (RP 2009:345). / The brain is at the centre of the nervous system, receiving and sending messages to control all parts of the body (RP 2009:345).
Synonyms: akaaka, pūnaha ioio, io, uaua, maihara
